The suspected mastermind of the jihadist cell dismantled in Belgium is still at large, a Belgian minister has said.
The comments from Justice Minister Koen Geens come amid reports the suspect is in Greece.
When asked if the ringleader remained on the run after four people were arrested in Athens on Saturday, Mr Geens told VRT television: "That is indeed the case."
He added: "Last night's arrests did not succeed in nabbing the right person. We are still actively looking for him and I presume we will succeed."
Belgian media have named the suspected leader of the cell uncovered by police in the eastern town of Verviers as Abdelhamid Abaaoud, a 27-year-old Belgian of Moroccan origin.
Two suspects were shot dead in a fierce gun battle with police during the raid, which smashed a cell plotting to kill Belgian officers on the street, local authorities said.
According to Belgian media, Abaaoud spent time fighting alongside Islamic State militants in Syria.
He was already known to security forces after appearing in an Islamic State video, at the wheel of a car transporting mutilated bodies to a mass grave.
Belgium's Flemish-language VTM channel said Abaaoud had made calls from Greece to the brother of one of the two heavily-armed suspects killed in Verviers.
A Greek police source said anti-terrorism investigators sent DNA samples and fingerprints to Belgium to establish whether Abaaoud was among the suspects arrested in Athens.
Initial reports had put the number of suspects in Greek custody at four but a police source said only two people were detained, at least one of whom was released without charge.
A spokesman for the Belgian federal prosecutor's office, Eric Van Der Sypt, said there was "no connection" between the suspects and their enquiry.
In Belgium, 13 people were arrested in connection with the probe. Five have been charged with "participating in the activities of a terrorist group."
Weapons, bomb-making materials, police uniforms and fake documents were found during searches of their homes.
Furthers searches were made Sunday in the Brussels district of Molenbeek where Abaaoud lived, media reports said.
Two fugitives who left Belgium immediately after the attack were arrested in France.
The Paris atrocities, in which 17 people were killed, have rekindled fears about young Europeans returning home after fighting alongside extremist groups in the Middle East.
Two of the three terrorists responsible for the attacks, brothers Said and Cherif Kouachi, were buried over the weekend in anonymous graves.
There has been no word on burial plans for the third gunman Amedy Coulibaly, who murdered five people before he was shot dead by police.
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